Ecology of small mammals and their ectoparasites in Mount Meru, Tanzania: insights for biodiversity conservation

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Date

2024-05

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Sokoine University of Agriculture

Abstract

This Thesis has been prepared in accordance with the Publishable Manuscripts format of the Sokoine University of Agriculture. The study on the elevational distribution of non-volant small mammals over time, their demographic characteristics, and their associations with arthropod ectoparasites was conducted between 2021 and 2022 at different elevations and habitats in Tanzania's second- highest mountain, Mount Meru. Mount Meru is one of the biodiversity hotspots in eastern Africa, hosting high level of species richness and endemism, and providing important ecosystem services. However, relatively, little is known about the ecology of small mammals and their arthropod ectoparasites. Similarly, little is known about the effects of environmental factors on the small mammal distribution, which potentially extending to the parasites they host. Tanzania is among the handful of countries in Africa where active foci of deadly zoonotic diseases that can be transmitted by small mammals and their ectoparasite vectors have been persistent. Moreover, Mount Meru is home for endemic and threatened species that are documented on the IUCN Red List. However, there is currently no available information regarding the possible role of arthropod ectoparasites of small mammals in the transmission of zoonotic disease. The over all goal of the study was to understand the ecological dynamics and distribution patterns of small mammals and their arthropod ectoparasites across different elevations of Mount Meru in Tanzania. The study aimed at addressing three key specific objectives: i) Assesing a decade-long change in the elevational distribution of non-volant small mammals on Mount Meru; ii) determining the breeding and demographic aspects of montemys delectorum, an east african endemic rodent in mount Meru; iii) Examining arthropod ectoparasites of two rodent species occurring in varied elevations on Tanzania’s second highest mountain. To address the first objective, the small mammals were surveyed from February to November 2021 at six elevations between 1500 m and 3650 m using Sherman and Havahart traps. The results show that two shrew and thirteen rodent species were recorded, including Crocidura newmarki and Lophuromys verhageni, which are endemic to the study area. Species richness was highest at mid-altitudes, in line with the general patterns globally, though less pronounced in Africa. The species Arvicanthis niloticus, Mastomys natalensis, Lemniscomys striatus, Dasymys incomtus, and Cricetomys ansorgei were not documented in 2009 but have been recorded in 2021. Upward shifts in species’ ranges were observed between 2009 and 2021, suggesting the influence of climate change on the small mammal community. It is important to note that the rodent species mentioned in Chapter 2 as Praomys species have been identified as Montemys delectorum after genetic identification and recent taxonomic revisions. For the second objective, a Capture – mark – release studies were undertaken in dense and open patchy forests in four 70 m X 70 m grids from February 2021 to June 2022. The findings revealed that the species exhibited bimodal patterns of breeding seasonality, characterized by a low rate occurring during the short interval between December and February and a pronounced rate taking place from May to July. The population of Montemys delectorum was unstable in the different forest types. There was a larger population in the patchy open forest (F = 8.1; df = 1; p = 0.0001) than in dense forest, suggesting its importance as a refuge for the species. To achieve the third objective, trapping of the two most dominant rodent species of the study area, Montemys delectorum and Rhabdomys dilectus, from five elevations (1500 to 3500 m), were examined for arthropod ectoparasite infestations. The results revealed that Ctenophthalmus calceatus cabirus, Dinopsyllus ellobius, Varroa rindereri, and Xenopsylla cheopis were recovered from the rodents. This study demonstrates that the elevational distribution of arthropod ectoparasites is influenced by host sex, host species, and environmental temperature. With rising altitude, parasite prevalence and abundance decrease, indicating that colder environmental conditions at higher altitudes inhibit parasite reproduction and development. This indicates conservation efforts should focus on higher altitudes. Moreover, the rodents and ectoparasites reported in this study have been identified as possible vectors of medical and veterinary diseases in different parts of Africa, including Tanzania. It is crucial to take precautions in order to effectively reduce the risk of parasitic infection, which can pose an additional threat to both the endemic and threatened species listed on the IUCN Red List as well as the overall biodiversity of the study area. Furthermore, molecular results from this study reported a previously unidentified mite species within the Mesostigmata taxonomic group that had previously been recognized exclusively as a parasite of honeybees. Further research efforts may provide insights into the significance of this particular species of mite, Varroa rindereri. Generally, while there remains much to be learned, this particular study establishes the foundation for future initiatives in wildlife disease surveillance and biodiversity conservation management. In abroader sense, the shift in the distribution of small mammals and their ectoparasites on Mount Meru is a microcosm of the broader challenges facing mountain ecosystems worldwide. These insights go beyond the immediate context of Mount Meru and suggest that we need to consider the long-term consequences of environmental dynamics on biodiversity conservation. As these complexities continue to shape mountain landscapes, their impact is not limited to species interactions but also touches the field of human well-being. The conservation of biodiversity is intrinsically linked to our own well- being, as biodiversity and human societies are intricately intertwined. Therefore, the findings of this research serve as an urgent reminder for effective conservation action.

Description

PhD Thesis

Keywords

Ecology, Small Mammals, Their Ectoparasites, Mount Meru, Biodiversity Conservation, Tanzania

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